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  • head medium wide
    barbells: lip - short filament + 5-6; chin 17-20
    face rim medium wide, undulating, serrated, end - flat triangular wing, rear edge forward
    rostral plates medium, < snout
  • head broad, rim wide, ends in strong spine
    rostral plates short, broad
    large nasal spine
    barbels: lip 1 v long, chin 31-42
  • face rim: wide, rippled edge, ends in wide, blunt triangle
    rostral plates short, broad, triangular
    Ds & pectoral: outer 1/2 black
    barbells: lip - 1 v long + 5; chin 11-13
  • head broad; rim wide, undulating, ends in wide wing
    rostral plates v short
    barbels: lip - 1 v long filament + 2, slender, chin - 8-9 in 4-5 clusters (each with 1 large)
    nostrils: strong curved spines
    pectoral: dark upper base, outer 2/3 black
    C: base dark bar, top 1/2 outer 1/3 black
  • head v broad, wide between eyes
    face rim not wide, end - sharp, flat spine
    rostral plates v short, broad
    barbells: lip - short filament + 5-12; chin - 12-18