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  • head shield few granules
    D spine > pectoral spine
    fins & barbels pale
    gill rakers: 1st arch 18-21, 2nd 17-21
    rear of pectoral spine - inconspicuous serrations
    A 18-21
  • tip snout to D: 39-40.7% SL
    snout 9.3-11.6% SL
    D2 dark
    gill rakers: 1st arch 14-16, 2nd 13-16
    fleshy papillae between gill rakers
  • D spine = pectoral spine
    fins & barbels pale
    gill rakers: 1st arch 19-20, 2nd 16-19
    rear of pectoral spine - inconspicuous serrations
    A 18-21
  • tip snout to D origin: 33.1-38%
    snout 6-8.6% SL
    fins dark, especially edges
    gill rakers: 1st arch 14-16, 2nd 13-16
    fleshy papillae between gill rakers
    A 21
  • body elongate
    head shield -dense granules
    bone behind head shield 8.2-9.8% SL, profile straight
    D spine > pectoral spine
    pectoral spine inconspicuous serrations
    not yellow
    top barbell reaches past pectoral spine base
    gill rakers: 1st arch 21-24, 2nd 19-22
    A 21-25
  • head 22.3-23% SL
    pelvics & pectorals intensely black
    gill rakers: 1st arch 17-18, 2nd 16-18
    pectoral spine short, long serrations
    A 19-21
  • body elongate
    head shield: dense granules
    D spine > pectoral spine
    pectoral spine small serrations
    not yellow
    bone behind head shield 11.7-18.9% SL, profile straight
    eye large (4.3-6.6% SL)
    top barbel usually passes pectoral spine
    gill rakers: 1st arch 18-23, 2nd arch 16-19
    A 20-24