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  • snout length ~1/4 disc width
    disc top covered with denticles
    tail: > disc, 1 row thorns
  • disc front rounded to ~straight
    top covered with denticles
    1 pair tubercles on shoulders
    tail long, no D, with low ventral keel
    tail spines well to rear
  • densely spotted
    tail < disc
  • snout length ~1/3 disc width
    disc top ~no denticles
    tail: > disc, no row thorns