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  • top head profile convex
    snout not upturned
    male C top edge smooth
    top tooth plates rough
    A present
  • snout short, conical
    C top and bottom lobes similar
    A present
  • top head profile convex
    snout upturned
    male: top edge of C smooth
    top tooth plates rough
    No A
  • snout short, conical
    C lobes similar
    A absent
  • top head profile straight
    snout straight
    male: top C edge + large tubercles
    top tooth plates smooth
    no A