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  • sucker small, weak, lacks papillae
  • head wide, body short
    top lip narrow
    incisors compressed, lower serrated
    sucker small, papillae patches: front, center, rear
    lower ~5 pectoral rays smaller
    pectoral base - small pad
  • head wide
    top lip wide, wider in center
    incisors rounded
    sucker large, papillae patches: front, center (0-2), rear
    pectoral base - small pad
  • wide head
    top lip wide, wider in center
    sucker large, papilla patches: front, sides, rear, 2 (close-set) central
    pectoral base: large fleshy pad
  • head narrow
    top lip wide, wider in center
    sucker large, papilla patches: front, center (V-shaped), rear
    pectoral base - small pad
  • head broad
    top lip wide, wider in center
    sucker papilla patches: front, sides, rear, 2 (wide-spaced) central
    pectoral base: large pad
  • head narrow
    sucker papilla: front & rear only
    pectoral base: small pad