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  • head deep, square; prominent crest with knob on nape
    operculum with blunt flap
    end top lip - barbel
    preopercle serrated
  • body quadrangular, robust
    low crest on nape
    D 23-27
    preoperculum serrated
    operculum: blunt spine
    A I-II, 20-26
  • very elongate
    no nape ridge
    D 54-60
    preoperculum smooth
    operculum: sharp spine