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  • tail > disc
    C: top lobe - height < 1/6 length, not joining bottom lobe
  • disc rhomboidal, corners angular
    disc +/- central row thorns
    tail: base broad, depressed, spiny, + large tubercles
    mouth broad
  • disc oval/circular, corners rounded
    tail base narrow, circular
    mouth narrow
    disc +/- central row thorns
  • disc robust, rhomboidal, corners angular/broadly rounded
    disc + central row thorns
    tail: base not wide, no flat, no tubercles
  • tail < disc
    C: top lobe - height ~1/4 length, joins bottom lobe