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disc ~rhomboidal/rounded
snout short to medium, tip stiff
eyes + semicircle of thornscenter disc: row thorns shoulder to C
nape thorns in triangle
top disc & tail : denticles coarse, not densetop dark/light grey, bottom darker
tail thorn rows: central & sidetail 60% TL -
disc heart shaped, round outer corners
snout short, blunt, rounded, tip stiff
eyes + semicircle of thorns1 row thorns shoulder to tail base
nape thorns in triangle
top disc & tail : denticles coarse, not densetop dark grey, bottom black
tail thorn rows: central & sidetail 56-65% TL -
disc heart shaped, outer corners rounded
snout medium length, angular, tip stiff
eyes + semicircle of thorns1 row thorns shoulder to tail
nape thorns in triangle
top disc & tail : denticles dense, finetop & bottom black
tail thorn rows: central & sidetail 53% TL