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  • disc heart-shaped, front edge undulating
    snout flexible, short, blunt
    thorns: 3 rows nape to D
    top disc: 4-8 symmetric white spots
    pelvic: front lobe 67-76% rear lobe
  • disc heart-shaped, front edge undulating
    snout flexible, short, blunt, pale underneath
    1 central interrupted row thorns: nape to D
    top disc mottled + ocelli
    pelvic: front lobe 65-80% rear lobe
  • disc heart-shaped, front edge undulating
    snout flexible, short, blunt, dark underneath
    3 central rows thorns: nape to D
    top disc: uniform to mottled
    pelvic: front lobe 66-85% rear lobe
  • disc heart-shaped, front edge undulating
    snout flexible, short, blunt
    3 central rows thorns: nape to D
    C: with lower lobe
    under disc dark
    lower jaws with central spatula
    pelvic: front lobe 65-80% rear lobe
  • disc heart-shaped, front edge undulating
    snout flexible, short, blunt
    3 central rows thorns, nape to D
    C without lower lobe
    under disc dark
    lower jaws: no central spatula
    pelvic: front lobe 66-77% rear lobe