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  • D1: black blotch at rear, no long spines
    D2 margin convex in male, straight in female
    opercular spine - 2 points, curved up
    back: irregular yellow bars
    A: black stripe in both sexes
  • head long
    D1: black rear spot; male - spine I filamentous
    D2 edge concave in both sexes
    opercular spine - 2 points, curved up
    A black stripe in female only
  • head short
    D1: no spot in adult; male: 1st spine filamentous
    D2 edge concave in both sexes
    side: 2 wavy yellow lines
    opercular spine - 2 points, curved up
    A stripe: female black, male red